North African mtDNA derived from Iberian one

Found at Dienekes' Anthropology Blog, original paper at DOI: 10.1353 (paywall).

MtDNA H lineages from Tunisia are less diverse and within the variability found in Iberia. Therefore they are most probably a derivate. This fits terribly well with what I have been pondering in the last months or even years about the early origins of North Africans and specifically of Oranian culture (also known as Iberomaurusian) , so I'm quite excited about it.

Let's reconsider all elements:

Genetics:

• North African mtDNA H derived from Iberian H (also notice the relatively high concentration haplogroup V in Tunisia and nearby areas, that must be of European origin as well)
• North African mtDNA U6 less diverse than Iberian one. The lack of U6 elswhere in Europe and the greater diversity of its derived subclade U6a in NE Africa, has led some scholars to think it arrived from West Asia. But overall Iberia has by large the highest diversity of this clade, followed by Morocco, including haplogroups U6b and U6c, that are not found in NE Africa. See my earlier post on U6 and the Maca-Meyer paper on the matter.
• Odd rather common R1b ill-studied clades in NE Africa (Sudan, Upper Egypt) and also in Northern Cameroon, where it's dominant among some groups. While in this case the diversity argument is not so clear (most Iberian and European R1b belongs to a single subclade - but not all), we can't forget that Y-DNA is potentially much more susceptible to drift and that, in Western Europe, was affected by the LGM bottleneck and the Epipaleolithic demographic movements after it may have spread into Africa. R1b is not dominant but it's still a somewhat important haplogroup in NW Africa (I understand that the Capsian/Afroasiatic countertide replaced it largely by E1b1). Most North African R1b haplotypes connect much better with European than with West Asian clades in fact (there is one exception though).

Archaeology:

• The curious synchretic SE Iberian Gravetto-Solutrean culture dates from c. 22,000 BP (late 20th century calibrations, today it'd be probably somewhat older, like 25,000 BP maybe). The Solutrean of Mallaetes and Parpalló is among the oldest ones (only surpassed by that of Dordogne) but, unlike what happened in the Franco-Cantabrian region, where it became dominant soon after, in SE Iberia, it suffered a Gravettizing reaction that created a unique techno-cultural complex. Some of their artifacts fit extremely well with the back-tipped style found in North Africa, that also follow the all-covering Solutrean style of retouch.
• The Oranian culture of North Africa, concentrated along the coasts, was early on called Iberomaurusian because the affinities with Iberian techno-complex appeared evident. Later, as doubts about its origin mounted up, it was renamed Oranian. In the last times it has been common to claim that Oranian arrived from Sudan or Egypt but that is not the least clear in fact. What does appear to have migrated from that area is the Epipaleolithic, maybe even Mesolithic (grain-gathering) Capsian culture, that has a more interior distributon and that was probably the one spreading Afroasiatic (Berber) language in the area. Oranian earliest dates are of c. 20,000 BP and its human remains are considered Crô-Magnon type, a type that was most common in Europe with the Gravettian culture (though in Mediterranean Iberia also within the "Solutrean").
  
• The Qurta rock art of Upper Egypt (right in the crossroads where later Capsian may have originated) is incredibly similar to European rock art, specially to that of Côa valley in Portugal (see previous post on this matter). This artwork is dated to c. 16,000 BP.
  

All these archaeological elements fit in a chronological sequence Iberia-NW Africa-NE Africa that would be partly reversed later on with the Capsian (which did not arrive to Iberia though). The rather good match with genetic identifiers also seems to support this scheme. Maybe some corners need to be polished but I am every day much more persuaded that the ultimate origin of North Africans is in Europe, specifically in Iberia, rather that West Asia - even if later waves from the Nile and West Asia itself may have obscured this correlation.
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Middle Paleolithic remains found at Basque cave

The research at Arlanpe cave (Lemoa, Biscay, Basque Country) has unveiled important findings from the early Middle Paleolithic period, remains dated to 300-150,000 BP.

Other remains found at this little known cave are from the late Solutrean (c. 17,000 BP) and as recent as the Roman period. Animal remains are also relevant findings, among which has special importance those of brown bear (Ursus arctos) because their genetic analysis have shown that they are more related to Eastern European bear than to the Alpine and Balcanic population.

Sources (all in Spanish):
- Harpea Kultur Elkartea: Arlanpe Project (with images of the lithic assamblages found)
- Gara
- EITB
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European Paleolithic geography

So where is Isturitz, what is Magdalenian, what the heck are you talking about?

Ok, let's recapitulate:

1. A week ago, I commented on Aurignacian origins. Aurignacian culture is believed to be the first manifestation of Homo sapiens (we and our ancestors) and it's known to have penetrated pretty fast in most of Europe, displacing the former inhabitants, the Neanderthals. I also posted a map by A. Mellars:



Aurignacians began experimenting with rudimentary art and eventually also colonized, even if sparsely, the Mediterranean shores of the Iberian peninsula.

2. Haven't yet posted on it, I think, but the next thing to happen, many thousand years later was the developement or arrival of Gravettian culture. Gravettian technology shows some correlation with Neanderthal-made Chatelperronian but it's also true that similar technologies were in use among West Asians and possibly other peoples. Earlier, when Paleolithic research was concentrated in France, it was thought that Chatelperronian and Gravettian were two phases of the same cuture (Perigordian) but this is not sustained anymore and now it's believed that Gravettian culture evolved in Central-Eastern Europe or even arrived from outside the continent (though there's lack of evidence for this yet). It is likely that the Crô-Magnon type, strictu sensu, is associated with this culture.

Gravettian is also a pan-European culture and replaced Aurignacian everywhere, even if in many places it looks intrusive (older techniques remain). In fact it reached further than Aurignacian, incorportaing Eastern Europe and even penetrating into the Caucasus and Zagros areas (West Asia) and with connections as far east as the Altai. With Gravettian art became something much more common. Some of the most fascinating examples are the venus figurines, like the one of Lespuges:

3. In Western Europe, Gravettian didn't last long alone. A new techno-culture appeared locally, the Solutrean and replaced Gravettian rather fast, specially in the Franco-Cantabrian region. Instead, in Mediterranean Iberia, Gravettian showed to be strong enough and survived in a hybrid form with Solutrean (Iberian Gravetto-Solutrean), that is probably at the origin of North-African Ibero-Maurusian culture later on.

Solutrean was therefore primarily a culture of the Franco-Cantabrian region:

The Franco-Cantabrian region
(red dots are main sites of Paleolithic art, light green are areas that were over sea level in the past, white shows glacial areas).

Gravettian also survived in Italy and Eastern Europe. Central Europe is believed by many to have been too cold to be inhabited, at least densely, in this time, as the Solutrean phase is coincident with the Last Glacial Maximum, some 20,000 years ago. Another area that was affected by this episode was the Zagros region of West Asia, where human inhabitation probably became impossible or very hazardous then.

Solutrean peoples were very advanced for their time, knowing the needle, the fishing hook and maybe even the bow and arrow. They also advanced quite a bit in art expression.

4. After some milennia, Solutrean evolved into Magdalenian locally in the Franco-Cantabrian region. While many sites with transitional industries are known, there are some questions still to answer because the Magdalenian is surprisingly similar to old Aurignacian, even if much more refined. Since c. 13,000 BP, as the greatest cold went away, Magdalenian expanded into Central Europe. It also shows signs of late expansion into Mediterranean Iberia but with a very marked Gravetization anyhow (Parpalloan) and lacking the characteristic bone tools of this industry elsewhere.

Magdalenian is, of course, best known for its magnificient mural art, though these peoples also made potable art, like their predecessors.

Horse from Lascaux cave (Franco-Cantabrian region)

And here is where the Isturitz findings fit in: Magdalenian, Franco-Cantabrian, bone tools... whale bone tools! Makes sense, right?

Where is Isturitz? Quick graphic answer:

Is that better? Hope so.

Another interesting map is this one of European Paleolithic art:

Red: mural art. Green: portable art.
Cyan thick lines: glacial maximum: areas under ice. Blue lines: ancient coastline.

It helps getting an idea of where did humans dwelt in the Upper Paleolithic. Yet the Mediterranean, even if not so dedicated to art as other areas, also had some population density probably.